Publications

Peer-Reviewed Publications

71 in 25+ years; 54 since arriving at HSU; student co-authors underlined.

2011

Colwell, M.A., J.J. Meyer, M.A. Hardy, S. McAllister, A. Transou, R. LeValley, & S.J. Dinsmore. Western Snowy Plovers Charadrius alexandrinus nivosus select nesting substrates that enhance egg crypsis and improve nest survival. Ibis 153:303-311.
Brindock, K.M. & M.A. Colwell. Habitat Selection by Western Snowy Plovers During the Nonbreeding Season. The Journal of Wildlife Management 75(4):786-793.

2010

Colwell, M.A. Shorebird Ecology, Conservation and Management. University of California Press.
Colwell, M.A., N.S. Burrell, M.A. Hardy, K. Kayano, J.J. Muir, W.J. Pearson, S.A. Peterson, & K.A. Sesser. Arrival times, laying dates, and reproductive success of Snowy Plovers in two habitats in coastal northern California. Journal of Field Ornithology 81:349-360.
Muir, J.J., & M.A. Colwell. Snowy Plovers select open habitat for courtship scrapes and nests. Condor 112:507-510.
Mullin, S.M., M.A. Colwell, S.A. McAllister, & S.J. Dinsmore. Apparent survival and population growth of Snowy Plovers in coastal northern California.

Wilson, C.A. & M.A. Colwell. Movements and Fledging Success of Snowy Plover(Charadrius alexandrinus ) Chicks. Waterbirds 33(3): 331-340.

2008

Bluso-Demers, J.D., M.A. Colwell, J.Y. Takekawa, & J.T. Ackerman. Space use by Forster 's terns breeding in South San Francisco Bay. Waterbirds 31:357-364.
Conklin, J.R., M.A. Colwell, & N.W. Fox-Fernandez. High variation in roost use by Dunlin wintering in California: Implications for habitat limitation. Bird Conservation International 18:275-291.
Conklin, J.R. & M.A. Colwell. Individual associations in a wintering shorebird population: do Dunlin have friends? Journal of Field Ornithology. 79:32-40.
Demers, S.A., M.A. Colwell, J.Y. Takekawa, & J.T Ackerman. Breeding stage influences space use of female American avocets in San Francisco Bay, California. Waterbirds 31:365-371.
Hardy, M.A. & M.A. Colwell. The impact of predator exclosures on Snowy Plover nesting success: a seven-year study. Wader Study Group Bulletin 115:161-166.

2007

Colwell, M.A., S.J. Hurley, J.N. Hall, & S.J. Dinsmore. Age-related survival and behavior of Snowy Plovers chicks. Condor 109:638-647.
Colwell, M.A., S.E. McAllister, C.B. Millett, A.N. Transou, S.M. Mullin, & Z.J. Nelson, C.A. Wilson, R.R. LeValley. Philopatry and natal dispersal of the Western Snowy Plover. Wilson Journal of Ornithology. 119:378-385.
Conklin, J.R. & M.A. Colwell. Interactions of predators and manmade object causes potentially significant mortality in a wintering shorebird population. Wader Study Group Bulletin 112:57-59.
Conklin, J.R. & M.A. Colwell. Diurnal and nocturnal roost site fidelity of Dunlin (Calidris alpina pacifica) at Humboldt Bay, California. Auk 124:677-689.

2006

Colwell, M.A. Egg-laying intervals in shorebirds: Influences of time, energy and risk of clutch loss. Wader Study Group Bulletin 111:50-59.
Mathis, R.L., M.A. Colwell, L.W. Leeman, & T.S. Leeman. Long-billed Curlew distributions in intertidal habitats: Scale-dependent patterns. Western Birds. 37:156-168.

2005

Connolly, L.M., & M.A. Colwell. Comparative use of longline oysterbeds and adjacent tidal flats by waterbirds. Bird Conservation International 15:237-255.
Colwell, M.A., C.B. Millett, J.J. Meyer, J.N. Hall, S.J. Hurley, S.E. McAllister, A.N. Transou, & R.R. LeValley. Snowy Plover reproductive success in beach and river habitats. Journal of Field Ornithology 76:373-382.
Leeman, T.S., & M.A. Colwell. Coastal pasture use by Long-billed Curlew at the northern extent of their non-breeding range. Journal Field Ornithology 76:33-39.

View 2004 - 1986 publications »

2004

Roth, J.E., J.P Kelly, W.J. Sydeman, & M.A. Colwell. Sex differences in space use of breeding Common Ravens (Corvus corax) in Western Marin County, California. Condor 106:529-539.
Moore, J.E., M.A. Colwell, R.L. Mathis, & J.M Black. Staging of Pacific flyway black brant in relation to eelgrass with special consideration for Humboldt Bay. Biological Conservation 115:475-486.

2003

Colwell, M.A., T. Danufsky, N. Fox-Fernandez, J.E. Roth & J.R Conklin. Variation in shorebird use of diurnal high-tide roosts: How consistently are roosts used? Waterbirds 26:484-493.
Danufsky, T. & M.A. Colwell. Winter shorebird communities and tidal flat characteristics at Humboldt Bay, California. Condor 105:117-129.

2002

Colwell, M.A., R.L. Mathis, L.W. Leeman, & T.S. Leeman. Space use and diet of territorial Long-billed Curlews (Numenius americanus) during the non-breeding season. Northwestern Naturalist 83:47-56.
Taft, O.W., M.A. Colwell, C.R. Isola, & R.J. Safran. Waterbird responses to experimental drawdown: implications for the multispecies management of wetland mosaics. Journal of Applied Ecology 39:987-1001.

2001

Leeman, L.W., M.A. Colwell, T.S. Leeman, & R.L. Mathis. Diets, energy intake, and kleptoparasitism of non-breeding Long-billed Curlews in a northern California estuary. Wilson Bulletin 113:194-201.
Colwell, M.A., T. Danufsky, R.L. Mathis, & S.W. Harris. Historical changes in the abundance and distribution of the American Avocet at the northern limit of its winter range. Western Birds 32:1-15.
Colwell, M.A. & R.L. Mathis. Seasonal variation in territory occupancy of non-breeding Long-billed Curlews in intertidal habitats. Waterbirds 24:208-216.

2000

Colwell, M.A. A review of territoriality in non-breeding shorebirds. Wader Study Group Bulletin 93:58-66.
Colwell, M.A. & K.R. Sundeen. Shorebird distributions on ocean beaches of northern California. Journal of Field Ornithology 71:1-15.
Isola, C.R., M.A. Colwell, R.J. Safran & O.W. Taft. Interspecific differences in habitat use by waterbirds foraging in managed wetlands of the northern San Joaquin Valley, California. Waterbirds 23:196-203.
Safran, R.J., M.A. Colwell, C.R. Isola & O.E Taft. Foraging site selection by nonbreeding white-faced ibis. Condor 102:211-215.
Colwell, M.A., & O.W. Taft. Waterbird communities in managed wetlands varying in depth. Waterbirds 23:45-55.

1999

Delehanty, D.J., R.C. Fleisher, M.A. Colwell, & L.W. Oring. Sex-role reversal and the absence of extra-pair fertilization in Wilson's phalarope. Animal Behavior 55:995-1002.

1998

Day, J. H., & M.A. Colwell. Waterbird communities in rice fields subjected to different farming practices. Colonial Waterbirds 21:185-197.
Dodd, S.L., & M.A. Colwell. Environmental correlates of nocturnal foraging by shorebirds at north Humboldt Bay, California. Wilson Bulletin 110:182-189.

1997

Colwell, M.A., A. Dubynin, A.Y. Koroliuk, & N.A. Sobolev. Russian nature reserves and conservation of biological diversity. Natural Areas Journal 17:56-68.
Colwell, M.A., & S.L. Dodd. Environmental and habitat correlates of pasture use by nonbreeding shorebirds. Condor 99:337-344.
Haig, S.M., C.L. Gratto-Trevor, T.D. Mullins, & M.A. Colwell. Population identification of western hemisphere shorebirds throughout the annual cycle. Molecular Ecology 6:413-427.
Safran, B.J., C.R. Isola, O.E. Williams, & M.A. Colwell. Benthic invertebrates and foraging locations of ten waterbird species in managed wetlands of the northern San Joaquin Valley, California. Wetlands 17:407-415.

1996

Dodd, S.L., & M.A. Colwell. Seasonal variation in nocturnal and diurnal distributions of nonbreeding shorebirds at North Humboldt Bay, California. Condor 98:196-207.

1995

Colwell, M.A. & S.L. Dodd. Waterbird communities and habitat relationships in Northern California. Conservation Biology 9:827-834.
Colwell, M.A., R.H. Gerstenberg, O.E. Williams, & M.G. Dodd. Four Marbled Godwits exceed the North American longevity record for scolopacids. Journal of Field Ornithology 66:181-183.

1994

Colwell, M.A. Shorebirds of Humboldt Bay, California: Abundance estimates and conservation implications. Western Birds 25:137-145.
Colwell, M.A. & J.R. Jehl, Jr. Wilson's Phalarope. In The Birds of North America, No. 83. (A. Poole, P. Stettenheim, and F. Gill, Eds.). Philadelphia: The Academy of Natural Sciences; Washington, DC: The American Ornithologists' Union.
Hunter, J.E. & M.A. Colwell. Ectoparasite (Phtheriptera) infestation of six shorebird species. Wilson Bulletin 94:400-403.

1993

Colwell, M.A. & R.J. Cooper. Estimates of coastal shorebird abundance: The importance of multiple counts. Journal of Field Ornithology 64:1-9.
Colwell, M.A. Shorebird community patterns in a seasonally dynamic estuary. Condor 95:94-103.
Colwell, M.A. & S.L. Landrum. Nonrandom shorebird distribution and fine-scale variation in prey abundance. Condor 95:104-114.

1992

Colwell, M.A. Wilson's phalarope nest success is not influenced by vegetation concealment. Condor 94:767-772.

1991

Oring, L.W., M.A. Colwell, & J.M. Reed. Lifetime reproductive success in the spotted sandpiper (Actitis macularia): sex differences and variance components. Behavioral Ecology & Sociobiology 28:425-432.
Oring, L.W., J.M. Reed, M.A. Colwell, D.B. Lank, & S.J. Maxson. Factors regulating annual mating success and reproductive success in spotted sandpipers (Actitis macularia). Behavioral Ecology & Sociobiology 28:433-442.

1990

Colwell, M.A. Effects of fluctuating water conditions on prairie shorebirds. Proceedings Prairie Conservation Endangered Species Conference 173-180.
Colwell, M.A. & L.W. Oring. Nest-site characteristics of prairie shorebirds. Canadian Journal of Zoology 68:297-302.

1989

Colwell, M.A. & L.W. Oring. Extra-pair mating in the spotted sandpiper: A female mate acquisition tactic. Animal Behavior 38:675-684.
Colwell, M.A. & L.W. Oring. Adult and natal philopatry of prairie shorebirds: Sex and species differences. Wader Study Group Bulletin 55:21-24.

1988

Colwell, M.A. & L.W. Oring. Habitat use by breeding & migrating shorebirds in southcentral Saskatchewan. Wilson Bulletin 100:554-566.
Colwell, M.A. & L.W. Oring. Breeding biology of Wilson's Phalarope (Phalaropus tricolor) in southcentral Saskatchewan. Wilson Bulletin 100:567-582.
Colwell, M.A. & L.W. Oring. Sex ratios and intrasexual competition for mates in a sex-role reversed shorebird, Wilson's phalarope (Phalaropus tricolor). Behavioral Ecology & Sociobiology 22:165-173.
Colwell, M.A. & L.W. Oring. Variable female mating tactics in a sex-role reversed shorebird, Wilson's Phalarope (Phalaropus tricolor). National Geographic Research 4:426-432.
Colwell, M.A., J.D. Reynolds, C.L. Gratto, D. Schamel, & D. Tracy. Phalarope philopatry. Proceeding 19th International Ornithological Congress, pp. 585-593.
Colwell, M.A., S.D. Fellows, & L.W. Oring. Chronology of shorebird migration at Last Mountain Lake National Wildlife Area, Saskatchewan, Canada. Wader Study Group Bulletin 52:18-22.
Colwell, M.A., C.L. Gratto, L.W. Oring, & A.J. Fivizzani. Effects of blood sampling on shorebirds: injuries, return rates, and clutch desertions. Condor 90:942-945.
Oring, L.W., A.J. Fivizzani, M.A. Colwell, & M.E. El Halawani. Hormonal changes associated with natural and manipulated incubation in the sex-role reversed Wilson's phalarope. General Comparative Endocrinology 72:247-256.
Colwell, M.A. & L.W. Oring. Wing fluttering display by incubating male Wilson's phalaropes. Canadian Journal of Zoology 66:2315-2317.

1987

Colwell, M.A. Seasonal shorebird abundance at Last Mountain Lake Wildlife Management Unit. Blue Jay 45:261-266.

1986

Colwell, M.A. The first documented case of polyandry for Wilson's Phalarope (Phalaropus tricolor). Auk 103:611-612.
Colwell, M.A. Intraspecific brood parasitism in three species of prairie-breeding shorebirds. Wilson Bulletin 98:473-475.
Reynolds, J.D., M.A. Colwell, & F. Cooke. Sexual selection and spring arrival times of red-necked and Wilson's phalaropes. Behavioral Ecology & Sociobiology 18:303-310.
Fivizzani, A.J., M.A. Colwell, & L.W. Oring. Plasma steroid levels in free-living Wilson's phalaropes. General Comparative Endocrinology 62:137-144.

Presentations at Professional Meetings

I regularly present and co-author papers at regional and national meetings. For example, in 2006-07, I co-authored 9 papers with students at three CA meetings: 1) Western Section of The Wildlife Society, Monterey; 2) Pacific Seabird Group, Asilomar; and 3) Society of Wetland Scientists, Sacramento.

Humboldt State University

wildlife

Mark Colwell, Ph.D.