1955: B.A.  Biology

Lawrence University, Appleton WI
   (Magna cum Laude, Phi Beta
    Kappa)

1957: M.S.  Fisheries

University of Michigan, Ann Arbor
   MI (Advisor: K.F. Lagler)

1961: Ph.D. Zoology/
   Limnology

University of Michigan, Ann Arbor
   MI (Advisor: G.H. Lauff)

1999-present

Senior Advisory Scientist, California Cooperative Fishery Research Unit and Adjunct Professor, Fisheries Dept., Humboldt State University, Arcata, CA

1994-1999

Distinguished Scientist, Ecosystem Restoration Department, South Florida Water Management District, West Palm Beach and Sanibel FL

1989-1993

Professor, Department of Biological Sciences and Director, Pymatuning Laboratory of Ecology, University of Pittsburgh, PA

1984-1989

Professor, Appalachian Environmental Laboratory, Center for Environmental and Marine Studies, University of Maryland, Frostburg, MD

1978-1984

Professor, Department of Fisheries and Wildlife and Department of Entomology, Oregon State University, Corvallis, OR

1972-1978

Professor, Department of Entomology, Department of Fisheries and Wildlife, Department of Zoology, and Kellogg Biological Station, Michigan State University, Hickory Corners, MI

1968-1972

Associate Professor, Department of Biology, University of Pittsburgh, Pittsburgh, PA

1961-1962

Assistant Professor, Department of Biology, Northwestern University, Northwestern University, Evanston, IL

1960-1961

Pre-/Post doctoral Instructor, Department of Zoology, University of
Michigan, Ann Arbor, MI

Carnegie Museum of Natural History, Pittsburgh PS (1994-present)

Florida Atlantic University, Department of Biological Sciences, Boca Raton, FL 1997-present

University of Florida, Institute of Food and Agricultural Science, Gainesville FL 1997-present

Florida International University, Department of Biological Science, Miami FL 1997-present

University of Miami, Rosenstiel School of Atmospheric and Marine Science, Miami FL 1998-present

University of South Florida, Department of Biology, Tampa FL 1999-present