• PhD, 1993, Harvard University
• MA, 1986, Harvard University
• BA, 1984, University of California, Santa Cruz

• Comparative Anatomy of the Vertebrates
• Introductory Zoology
• History of Biology
• California Natural History
• Electron Microscopy

Graduate Students

Former:

• 2024, Emily Gremling: "Comparative morphology and ultrastructure of olfactory epithelia in plethodontid salamanders: Effect of life history variation"
• 2024, Giuseppina (Sun) Lanzilli: "Comparing alternative developmental modes: Structure and gene expression in the olfactory system of plethodontid salamanders"2023, Jaclyn Patmore: "Olfactory organ morphology and ultrastructure of the larval Koh Tao Island caecilian (Ichthyophis kohtaoensis)"
• 2021, Kyle Orr: “The homology of sarcopterygian gills.”
• 2020, Ethan Snee: “An ecomorphological analysis of locomotion in larvae and neotenes of two salamander species: Dicamptodon tenebrosus (stream-type) and Ambystoma gracile (pond-type).”
• 2019, Adrian Macedo: "Life history of the coastal tailed frog (Ascaphus truei) across an elevational gradient."
• 2017, Christian Brown - Habitat use and movement patterns of two redwood forest salamanders, Aneides vagrans and Ensatina eschscholtzii, with an examination of the efficacy of PIT tags for marking small plethodontids
• 2016, Jennifer Tucker - Effectiveness of cutaneous antifungal microbe bioaugmentation for disease prevention in the Cascades Frog, Rana cascadae, of northern California
• 2015, Adam Cummings - Evolution of an aquatic courtship pheromone in the genus Taricha
• 2013, Timothy Girod - Bullfrog (Lithobates catesbeianus) diet composition, life history, and time period for larval development in California's north coast
• 2012, Michael Sun - Batrachochytrium dendrobatidis prevalence in Northern Red-legged Frogs (Rana aurora) - 10 years later
• 2011, Shannon Morago - A mechanism to increase chemosensory perception in humans?
• 2009, Larkin Chases - Olfactory response to food in larval Coastal Giant Salamanders, Dicamptodon tenebrosus
• 2008, Carrie Sendak - Movement patterns and habitat use of the Northern Red-legged Frog, Rana aurora
• 2006, Noelle Benzekri - Olfactory structure and metamorphosis in the Coastal Tailed Frog, Ascaphus truei
• 2004, Nathan Nieto - Chytridiomycosis in larval Red-legged Frogs in and near Redwood National Park
• 2004, Jeremy Stuelpnagel - Olfactory metamorphosis in the Coastal Giant Salamander, Dicamptodon tenebrosus
• 2003, Stein Coriell - Environmental factors affecting paedomorphosis in the Coastal Giant Salamander
• 2001, Jason Lowe - Population structure of two sympatric salamanders, Plethodon elongatus and Ensatina eschscholtzii